While founding wounded Villanelle, Eve is called back to London to investigate a murder case believed to have been ordered by the Twelve. Villanelle has a rough time as she makes her way from a Paris hospital to London.

Eve meets her new team and deduces that the murderer is not Villanelle, but a new assassin for the Twelve whom she nicknames “the Ghost”.

Villanelle is ordered by Raymond to perform a clean assassination like the Ghost. She kills her target in a lift and delivers lipstick to Eve with the name “Love in an Elevator” to ensure that Eve knows who performed the assassination.

Villanelle tries to get Eve’s attention with an assassination in Amsterdam based on The Corpses of the De Witt Brothers but is angered when Eve does not show up to investigate.

In a desperate attempt to get closer to Villanelle, Eve puts out a hit on herself, hiring Villanelle to do the job.

With new information about Aaron Peel, Villanelle goes undercover as a New Yorker named Billie to befriend Aaron’s sister, Amber but there is conflict between Villanelle and Aaron.

Aaron apologises to Villanelle and invites her to go to Rome. Villanelle interrogates Niko whilst Eve visits the psychologist. In Rome, Villanelle discovers Aaron’s plans but Eve becomes jealous of their relationship.